4 GV call(s) male advance with chips, 4 GV call(s) male advance without chips, 3 GV call(s) male advance, 5 GV call(s) female advance, 1 F call(s) "ghostly penguin", 1 G call(s) "ghostly penguin" and "penguin dance", 4 GV call(s) food chirps, 23 GV call(s) alarm chirps, 1 GV call(s) distress call, 2 GV call(s) barge trill, 6 GV call(s) aggressive chirps, 2 F call(s) whirring, 2 G call(s) rearing with whirring, 1 G call(s) copulation trill