Mixed-conifer stands of grand fir (Abies grandis (Dougl. ex D. Don) Lindl., Douglas-fir (Pseudotsuga menziesii var. glauca (Beissn.) Franco), and ponderosa pine (Pinus ponderosa Dougl. ex Laws.) were fertilized with nitrogen and combination treatments of nitrogen, phosphorus, potassium, and sulfur to test their effects on trees and associated insects during an outbreak of the western spruce budworm (Choristoneura occidentalis Freeman).