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The Viral Protein A238L Inhibits TNF-α Expression through a CBP/p300 Transcriptional Coactivators Pathway

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dc.contributor Ministerio de Educación y Ciencia (España)
dc.contributor Wellcome Trust
dc.contributor Fundación Ramón Areces
dc.creator Granja, Aitor G.
dc.creator Nogal París, María Luisa
dc.creator Hurtado, Carolina
dc.creator Aguila, Carmen del
dc.creator Carrascosa, Ángel L.
dc.creator Salas, María Luisa
dc.creator Fresno, Manuel
dc.creator Revilla Novella, Yolanda
dc.date 2008-06-09T14:13:57Z
dc.date 2008-06-09T14:13:57Z
dc.date 2006-01
dc.date.accessioned 2017-01-31T01:37:10Z
dc.date.available 2017-01-31T01:37:10Z
dc.identifier The Journal of Immunology, 2006, 176: 451-462
dc.identifier 0022-1767 (Print)
dc.identifier 1550-6606 (Online)
dc.identifier http://hdl.handle.net/10261/4921
dc.identifier.uri http://dspace.mediu.edu.my:8181/xmlui/handle/10261/4921
dc.description Article available at http://www.jimmunol.org/cgi/content/abstract/176/1/451
dc.description African swine fever virus (ASFV) is able to inhibit TNF--induced gene expression through the synthesis of A238L protein. This was shown by the use of deletion mutants lacking the A238L gene from the Vero cell-adapted Ba71V ASFV strain and from the virulent isolate E70. To further analyze the molecular mechanism by which the viral gene controls TNF-, we have used Jurkat cells stably transfected with the viral gene to identify the TNF- regulatory elements involved in the induction of the gene after stimulation with PMA and calcium ionophore. We have thus identified the cAMP-responsive element and 3 sites on the TNF- promoter as the responsible of the gene activation, and demonstrate that A238L inhibits TNF- expression through these DNA binding sites. This inhibition was partially reverted by overexpression of the transcriptional factors NF-AT, NF-B, and c-Jun. Furthermore, we present evidence that A238L inhibits the activation of TNF- by modulating NF-B, NF-AT, and c-Jun trans activation through a mechanism that involves CREB binding protein/p300 function, because overexpression of these transcriptional coactivators recovers TNF- promoter activity. In addition, we show that A238L is a nuclear protein that binds to the cyclic AMP-responsive element/3 complex, thus displacing the CREB binding protein/p300 coactivators. Taken together, these results establish a novel mechanism in the control of TNF- gene expression by a viral protein that could represent an efficient strategy used by ASFV to evade the innate immune response
dc.description This work was supported by grants from Ministerio de Educación y Ciencia (BFU2004-00298/BMC), the Wellcome Trust (075813/C/04/z), and by an institutional grant from the Fundación Ramón Areces. C.H. was a fellow from Fundación Ramón Areces.
dc.description Peer reviewed
dc.format 3436629 bytes
dc.format application/pdf
dc.language eng
dc.publisher American Association of Immunologists
dc.rights openAccess
dc.subject ASFV
dc.subject A238L protein
dc.title The Viral Protein A238L Inhibits TNF-α Expression through a CBP/p300 Transcriptional Coactivators Pathway
dc.type Artículo


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